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The earliest mammaliaforms are difficult to assess because the fossil record is poor and because their distinctive morphologies cannot be directly compared with more recent mammaliaforms. This is especially true for the haramiyid genus  Theroteinus , only known in the Saint-Nicolas-de-Port locality (Rhaetian, France). This study presents a new definition of the type-species  Theroteinus nikolai . A new species  Theroteinus rosieriensis , sp. nov., is named and distinguished by the lingual shift of distal cusps, a larger size, and a stockier occlusal outline. Comparisons with  Eleutherodon ,  Megaconus  and  Millsodon  suggest that  Theroteinus  has potential close relatives among the Jurassic haramiyids.

A reappraisal ofTheroteinus(Haramiyida, Mammaliaformes) from the Upper Triassic of Saint-Nicolas-de-Port (France)