A revision of the lower Pennsylvanian Alethopteris lonchitica (auctorum) and its identity with Alethopteris urophylla

One of the most common lower Pennsylvanian (upper Namurian, lower Westphalian) pteridosperms is a species commonly recorded as Aletho pteris lonchitica Schlotheim, 1820 ex Sternberg, 1825. However, the photographic illustration of the lectotype of Aletho pteris lonchitica by Zodrow & Cleal (1998) has allowed a clear distinction from Aletho pteris lonchitica (auctorum) which corresponds, in the main, to Aletho pteris urophylla (Brongniart, 1834) Göppert, 1836. This species is redescribed here, and its holotype is illustrated photographically for the first time. The published records, backed up by illustration, are analysed with reference to lists of synonymy. The geographic and stratigraphic distribution of Aletho pteris urophylla is discussed, and illustrations of this species are provided from Nova Scotia, Scotland, England, Wales, the North of France, and NW as well as SW Spain. Aletho pteris lonchitica was identified with Aletho pteris lonchitifolia Bertrand, 1932 by Zodrow and Cleal, but the present writers prefer an identification with Aletho pteris friedelii Bertrand, 1932. The corresponding taxonomic implications are discussed.