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E cosystem science and evolutionary biology have long been infrequent and uncomfortable bedfellows (Hagen 1992, Holt 1995, Mclntosh 1985). However, the convergence of a global decline in biodiversity and global alterations in biogeochemical cycles provides motivation to overcome past inhibitions. Currently, attempts are being made {Jones and Lawton 1995) to understand relationships between the foci of evolutionary biology (the individual in its species population) and ecosystem science (energy and material flow and storage). Analysis of relationships between species and ecosystems requires a framework appropriate for moving between levels

Organism Size, Life History, and N:P Stoichiometry