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age (5−10 V/cm) runs necessary and highlights the limitations of using Tris as a cation for DNA electrophoresis. Another component of currently used buffers, EDTA, is now largely superfluous, since most DNA samples are readily soluble and since commonly used enzymes today would not carry an undesirable enzymatic activity under electrophoretic conditions.

Sodium boric acid: a Tris-free, cooler conductive medium for DNA electrophoresis