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Previous studies have explored the relationship between termite branch tunnel geometry and foraging efficiency in a model simulation in which foraging efficiency,  γ,  for two termite species,  Coptotermes formosanus  Shiraki and  Reticulitermes flavipes  (Kollar) (Isoptera: Rhinotermitidae), was investigated in response to two variables, the probability of tunnel branching ( P branch  ) and the probability of tunnel branch termination ( P term). It was found that simulated tunnel patterns based on empirical data did not have maximum foraging efficiency. We hypothesized that termites could increase their foraging efficiency in response to landscape heterogeneity. The present study investigated how termites could control the two variables,  P branch   and  P term  , in response to the external environment in terms of tunnel network connectivity. It was found that the best simulated strategy for  C. formosanus  and  R. flavipes  termites would occur if both  P branch   and  P term   were increased together. This study provides possible mechanisms for foraging strategies in subterranean termites and a baseline for future empirical work.

A Constraint Condition for Foraging Strategy in Subterranean Termites