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A new inquiline oak gall wasp species, Saphonecrus yukawai Wachi, Ide &amp; Abe, sp. nov. (Hymenoptera: Cynipidae: Synergini), is described from Japan. Although most inquiline oak gall wasps are associated with cynipid galls on Quercus trees, the adult wasps of S. yukawai emerged from galls induced by the gall midge Ametrodiplosis acutissima (Monzen) (Diptera: Cecidomyiidae) on Quercus (sect. Cerris) acutissima Carruthers. Host shift from cynipid galls to cecidomyiid galls in inquiline oak gall wasps is discussed.

A New Inquiline Species ofSaphonecrus(Hymenoptera: Cynipidae: Synergini) Associated with Cecidomyiid Galls on Oak Trees in Japan