Generation of periodic travelling waves in cyclic populations by hostile boundaries

Many recent datasets on cyclic populations reveal spatial patterns with the form of periodic travelling waves (wavetrains). Mathematical modelling has identified a number of potential causes of this spatial organization, one of which is a hostile habitat boundary. In this paper, the author investigates the member of the periodic travelling wave family selected by such a boundary in models of reaction–diffusion type. Using a predator–prey model as a case study, the author presents numerical evidence that the wave generated by a hostile (zero-Dirichlet) boundary condition is the same as that generated by fixing the population densities at their coexistence steady-state levels. The author then presents analysis showing that the two waves are the same, in general, for oscillatory reaction–diffusion models with scalar diffusion close to Hopf bifurcation. This calculation yields a general formula for the amplitude, speed and wavelength of these waves. By combining this formula with established results on periodic travelling wave stability, the author presents a division of parameter space into regions in which a hostile boundary will generate periodic travelling waves, spatio-temporal disorder or a mixture of the two.