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1 Department of Chemistry, KAIST, Daejeon, 305-701, Korea 2 Division of Electron microscopic Research, Korea Basic Science Institute, Daejeon, 305-333, Korea 3 Department of Biological Sciences, KAIST, Daejeon, 305-701, Korea 4 Division of Magnetic Resonance, Korea Basic Science Institute, Cheongwon, Chungbuk, 363-883, Korea 5 Present address: Immunotherapy research center, Korea Research Institute of Bioscience and Biotechnology, Daejeon, 305-806, Korea 6 Present address: Medical Proteomics Research Center, Korea Research Institute of Bioscience and Biotechnology, Daejeon, 305-806, Korea

Srs2 possesses a non-canonical PIP box in front of its SBM for precise recognition of SUMOylated PCNA