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Valerio Sbragaglia*, Lorenzo Morroni Lorenzo Bramanti, Boris Weitzmann, Robert Arlinghaus, and Ernesto Azzurro Department of Biology and Ecology of Fishes, Leibniz-Institute of Freshwater Ecology and Inland Fisheries, Müggelseedamm 310, 12587 Berlin, Germany Institute for Environmental Protection and Research (ISPRA), Via del Cedro 38, 57122 Livorno, Italy Department of Life and Environmental Sciences, Polytechnic University of Marche, Via Brecce Bianche, 60131 Ancona, Italy Sorbonne Universitè, CNRS, Laboratoire d’Ecogéochimie des Environnements Benthiques (LECOB), Observatoire Oceanologique Banyuls sur mer. 1 Avenue Pierre Fabre, 66650 Banyuls sur mer, France Parc Natural del Montgrı́, les Illes Medes i el Baix Ter, L’Estartit, Catalunya, Spain Division of Integrative Fisheries Management, Department of Crop and Animal Sciences, Faculty of Life Sciences, Humboldt-Universität zu Berlin, Invalidenstrasse 42, 10115 Berlin, Germany Stazione Zoologica Anton Dohrn (SZN), Villa Comunale 80121 Napoli, Italy *Corresponding author: tel: þ39 0650074030; fax: þ39 0586896248; e-mail: valeriosbra@gmail.com. Equally contributed to this study.

Spearfishing modulates flight initiation distance of fishes: the effects of protection, individual size, and bearing a speargun