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S. L. Kram1*, N. N. Price1†, E. M. Donham2, M. D. Johnson1, E. L. A. Kelly1, S. L. Hamilton2, and J. E. Smith1 Center for Marine Biodiversity and Conservation, Marine Biology Research Division, Scripps Institution of Oceanography, La Jolla, CA 92093-0202, USA Moss Landing Marine Laboratories, 8272 Moss Landing Road, Moss Landing, CA 95039, USA *Corresponding author: tel: +415 497 5811; fax: +858 822 2167; e-mail: skram@ucsd.edu Present address: Bigelow Laboratory for Ocean Sciences, 60 Bigelow Drive, East Boothbay, ME 04544, USA.

Variable responses of temperate calcified and fleshy macroalgae to elevated pCO2 and warming