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This is a pre-copyedited, author-produced version of an article accepted for publication in Behavioral\ud Ecology following peer review. The version of record is available online at: doi: 10.1093/beheco/arv119. © The Author 2015. Published by Oxford University Press on behalf of the International Society for Behavioral Ecology. All rights reserved. For permissions, please e-mail: journals.permissions@oup.comBiotechnology and Biological Sciences Research Council (BBSRC

In search of the Darwinian Holy Trinity in cognitive evolution: a comment on Croston et al.