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Laurent Keller," Serge Aron, and Luc Passera Institut de Zoologie et d'Ecologie Animale, University of Lausanne, Batiment de Biologie, 1015 Lausanne, Switzerland, and Zoologisches Institut, Bern University, Ethologische Station Hasli, Wohlenstrasse 50a, 3032 Hinterkappelen, Switzerland, and ''Centre de Recherche en Biologie du Comportement (URA CNRS 664), Universite Paul-Sabatier, F 31062 Toulouse Cedex, France

Internest sex-ratio variation and male brood survival in the ant Pheidole pallidula