Carbon limitation, stem growth rate and the biomechanical cause of Corner’s rules

Corner's rules describe a global spectrum from large-leaved plants with thick, sparingly branched twigs with low-density stem tissues and thick piths to plants with thin, highly branched stems with high-density stem tissues and thin piths. The hypothesis was tested that, if similar crown areas fix similar amounts of carbon regardless of leaf size, then large-leaved species, with their distantly spaced leaves, require higher stem growth rates, lower stem tissue densities and stiffnesses, and therefore thicker twigs.