New Aspects in Floral Development of Papilionoideae: Initiated but Suppressed Bracteoles and Variable Initiation of Sepals

The increase of molecular data and the resulting insights into legume systematics make the search for new morphological characters and a careful re-investigation of already stated characters necessary. Bracteoles are small, reduced leaves borne close to the base of lateral branches. Although they seem unimportant in older buds, they have an ecological function in protecting the sepal primordia. Furthermore, a morphogenetic function in mediating the onset of sepal initiation is suspected in the literature. The occurrence of bracteoles varies within Papilionoideae, and their distribution is used in legume systematics. But this is open to criticism, because there is a tendency to use 'absent' for 'caducous'. Thus attention here was paid to the initiation of bracteoles as well as to the sequence of sepal initiation.