Coloration and structure of Taiwanese bronze scarab (Anomala expansa)

The structure and optical properties of the elytra of Taiwanese bronze scarabs (Anomala expansa) were investigated. The elytron is composed by a scale-like texture on the surface and a repeated bi-layers underneath. The reflectivity spectrum shows iridescence that the peaks shifts from yellow to green color regions as long as the incident angle (θ) increases. A shoulder peak at 600 nm (θ = 20o) is absent after gently scraping the surface, which indicates that the yellow color is contributed by the scale-like texture. The remaining peak at 550 nm (θ = 20o) in the spectrum resembles the simulated spectrum by assuming the refractive index of n = 1.54 and n = 1 for the adjacent bi-layered structure. This implies that the multi-layered structure contributes to the green coloration as well as the iridescent effect