Nowak et al. reply

Replying to: P. Abbot et al. 471, 10.1038/nature09831 (2011); J. J. Boomsma et al. 471, 10.1038/nature09832 (2011); J. E. Strassmann et al. 471, 10.1038/nature09833 (2011); R. Ferriere & R. E. Michod 471, 10.1038/nature09834 (2011); E. A. Herre & W. T. Wcislo 471, 10.1038/nature09835 (2011)Our paper challenges the dominant role of inclusive fitness theory in the study of social evolution. We show that inclusive fitness theory is not a constructive theory that allows a useful mathematical analysis of evolutionary processes. For studying the evolution of cooperation or eusociality we must instead rely on evolutionary game theory or population genetics. The authors of the five comments offer the usual defence of inclusive fitness theory, but do not take into account our new results.