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The role of the species scale as the Quantum Gravity (QG) cut-off has beenrecently emphasised in the context of the Swampland Program. Along these lines,we continue the quest for a global definition of the species scale within themoduli spaces of QG by exploiting duality symmetries, the known asymptoticbehaviours imposed by the presence of infinite towers of light states, as wellas the known behaviour of higher curvature corrections to the Einstein-Hilbertaction in certain String Theory set-ups. In those cases, we obtain a consistentresult for the identification of the (global) species scale as the QuantumGravity cut-off, but also present some puzzles related to the suppression ofhigher-dimensional corrections and certain minor ambiguities that may arise inthe deep interior of moduli space.

On the Species Scale, Modular Invariance and the Gravitational EFT expansion