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A flat-headed goby Luciogobius guttatus is distributed in the temperate coast and tidal zone of the Korean Peninsula, Japan, and the Northwest Pacifi c of China (Nelson, 1994). In general, the coastal region where such an intertidal fi sh lives has turbulent environments: a fluctuating water amount, a variety of salinity, and wave forces caused dramatically by periodic physical and chemical conditions (Horn et al., 1999). In particular, the periodic tidal cycle not only applies a physical effect directly to the breaking of boulders, the movement of gravels, the fl oating of sand and mud particles but also exposes the land of intertidal zone in the air during the low tide period (Van Straaten & Kuenen, 1958). Therefore, the intertidal fi shes exposed at the low tide are faced with a grave stress by physical impacts, a desiccant with emerging partial skin in tide pool and an underwater with low oxygen level (Congleton, 1980; Richards, 2011). To overcome these severe-repeated hard aquatic environments, they came to have special organs to perform various functions. Among them, the olfactory organ has been known as a good apparatus to recognize unstable and extreme environments. Olfaction in fi shes is a biologically essential chemoreception. It provokes important behaviors for the survival of fi sh such as feeding, reproduction, avoiding predator and intraspecifi c communication (Hara, 1975). In teleost, the olfactory organ responsible for this reception generally consists of peripheral organs, olfactory tract nerves and olfactory bulbs, conducting signal process and transduction with appropriate cells (Hara, 1986). It also varies in not only gross morphology, position, but also the number and morphology of the receptor cell, and the distributional pattern of the epithelium between species (Yamamoto, 1982). Kuciel et al. (2013) in the recent study reported that the olfactory organ of the intertidal fi shes

The Anatomy and Histoarchitecture of the Olfactory Organ in the Korean Flat-Headed Goby Luciogobius guttatus (Pisces; Gobiidae)