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The Oryctocephalus indicus biozone (Wuliuan, Miaolingian) is recognised in the Sumna Valley, in the southeastern part of the Spiti region in the Himalaya, based on the first appearance datum and last appearance datum of Oryctocephalus indicus. The biozone is approximately 5.6 m thick (17.8–23.4 m), and it contains the trilobites Oryctocephalus indicus, Pagetia significans and Kunmingaspis pervulgata. Comparative studies of the lithological variations across the Cambrian Series 2–Wuliuan (Miaolingian) transition in the Parahio and Sumna valleys (southeastern part of the Spiti region) show that the Wuliuan (Miaolingian) deposits transgressed over the undulatory surface of Cambrian Series 2. The uppermost part of the Cambrian Series 2 deposits in the southeastern part of the Spiti region is characterised by a reddish-brown ferruginous, very coarse grained sandstone unit, which indicates a diastem prior to the Wuliuan (Miaolingian) transgression.

Miaolingian transgression and the Oryctocephalus indicus biozone in the Sumna Valley (Spiti), Himalaya, India