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During mitosis, accurate segregation of the chromosomes duplicated at the S phase of the cell cycle relies on the successful of their proper attachment to microtubules of the mitotic spindle and their alignment at the metaphase plate before the onset of anaphase. Attachment errors or failure to align can lead to chromosome gains or losses, a condition known as chromosome instability (CIN) which is a common feature amongst many cancers.

The Kinetochore and Mitosis: Focus on the Regulation and Correction Mechanisms of Chromosome-to-Microtubule Attachments