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STRETCHING ACROSS the Pacific between 45°N and the coast of Alaska (Fig. 1) is an oceanic region referred to as subarctic. It is part of a broader oceanic region in which major phytoplankton nutrients (nitrate, phosphate, and silicate) apparently are never depleted from the surface layer. Persistently high nutrient concentrations are found along the entire eastern edge of the Pacific with westward extensions in the subarctic and along the equator and with eastward extensions through Drake Passage and then around the globe south of 35°S (Reid, 1962). Ecological relationships vary dramatically along this nutrient-replete belt, and assembly of a complete comparative ecology for all of its distinctive parts has only begun. Martin (1990; 1991, this issue) has opened a new chapter in this analysis by suggesting that limited availability of soluble iron may be a common factor preventing major nutrient depletion throughout the nutrient-replete belt. The initial observations leading to this suggestion (Martin and Fitzwater, 1988; Martin et al., 1989) were made in the subarctic Pacific, which immediately stimulated our interest as workers in the subarctic Pacific Ecosystem Research (SUPER) Program. Although the testing of Martin's hypothesis is still in an early phase (Banse, 1990; Martin et al., 1990), it is an attractive idea. It is one of several candidate explanations for the dominance of subarctic Pacific phytoplankton by cells of very small size. Given a flora composed of small cells, most other observations on subarctic Pacific ecology fit into a sensible, logical scheme. Perhaps similar schemes can be found for other sectors of the highnutrient belt. Iron limitation, if it proves impor-

Ecological Processes in the Subarctic Pacific: Iron-Limitation Cannot Be the Whole Story