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The twilight zone of the Earth’s carbon cycle The oceans’ mesopelagic layer or ‘Twilight Zone’ (200–1,000 m) acts as the gateway for long-term storage of atmospheric carbon in the deep ocean, mitigating global warming due to human carbon emissions. In the OPERA project, we will perform comprehensive model simulations, constrained by observations, to better understand and quantify mesopelagic carbon budgets (and their uncertainty) in the North Atlantic Ocean.