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Reproductive traits of female Rana nigromaculata inhabiting northern Ina Basin, Nagano Prefecture, were studied for two breeding seasons. The breeding season lasts at least two months between mid May and early July. This prolonged breeding is caused by markedly asynchronous development of ovarian eggs among females. Smaller females tend to breed later in the season than larger ones. Clutch size and ovum size are positively correlated with female snout-vent length.

Female Reproductive Traits in a Population of the Pond Frog, <i>Rana nigromaculata</i>, with Prolonged Breeding Season