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Rhopalosomatidae are an unusual family of wasps (Hymenoptera: Aculeata)\udcomprising less than 100 species found in the tropics and subtropics of\udall continents except Europe and Antarctica. Whereas some species resemble\udnocturnal Ichneumonidae, others might be mistaken for spider wasps or\uddifferent groups of brachypterous Hymenoptera. Despite their varied\udmorphology, all members of the family supposedly develop as larval\udectoparasitoids of crickets (Orthoptera: Grylloidea). Here, we report on the\udfirst record of a fossil rhopalosomatid larva which was discovered in\udmid-Cretaceous amber from northern Myanmar (Burma). The larva is attached to\udthe lateral side of a cricket between the metafemur and the abdomen,\udimpacting the natural position of the hind leg, exactly as documented for\udmodern species. Additionally, the larval gestalt is strikingly similar to\udthose of extant forms. These observations imply that this behavioral\udspecialization, e.g., host association and positioning on host, likely\udevolved in the stem of the family at least 100 million years ago

fossil record

The wasp larva's last supper: 100 million years of evolutionary stasis in the larval development of rhopalosomatid wasps (Hymenoptera: Rhopalosomatidae)