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A shallow Sargassum zone at around1to2m depth was followed sequentially by an Eisenia zone at2-7mdepth,a Dilophus zone less than1m wide,and a crustose coralline algal zone furthest off shore.Cobblestone areas were found here and there at2to3m depth.Samples handpicked and/or obtained by air-lifting were obtained around a station in each zone and a cobblestone area in June,August,October, and December,1990,and March,1991.A matrix of Pearson's correlation coefficients of species composi tion with respect to the analytical unit of•gsite•~time•hwas scaled multidimensionally.Behavioral areas of the MDS plots were similar between the Sargassum zone and cobblestone area,and between the Dilophus and crustose coralline algal zones,but the Eisenia zone was quite separate from these.A dendrogram of species-specific distribution pattern with respect to the analytical unit of•gspecies•~time•hshowed three strata differing in linkage levels,i.e.18subclusters with linkage levels of extremely low dissimilarity,nine clusters of low dissimilarity linkage,and three superclusters linked at relatively high levels of dissimilar ity.Out of the nine clusters,Clusters1,3,5,6,and8were characterized by having their highest relative densities in the crustose coralline algal zone,the Dilophus zone,the Eisenia zone,the Sargassum zone,and the cobblestone area,respectively.The respective superclusters were characterized by high relative densi ties in the following areas:Supercluster1,either the Dilophus zone or the crustose coralline algal zone; Supercluster2,the Eisenia zone;Supercluster3,either the Sargassum zone or the cobblestone area. Subclusters corresponded to subtle variations of cluster characteristics.These facts demonstrate the im portant role of algae in the structure of benthic animal distribution.

benthos research/nihon bentosu gakkaishi/nihon bentosu kenkyukaishi/nihon bentosu kenkyukai renrakushi

Distribution of Benthic Invertebrates in Relation to Zonal Structure of Algal Communities in a Rocky Sublittoral Area, Tomari-hama, Oshika Peninsula, Northern Japan