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Two varieties of Aucuba japonica differ in ways that can be considered adaptive to differing geo-climatic conditions intheir respective distribution ranges. Irrespective of growth stage, the mean leaf size of A. japonica var. japonica was significantlylarger than A. japonica var. borealis. Smaller leaf size and ultimately smaller stature of A. japonica var. borealisare an advantage under the higher snow load and lower temperatures in the forests along the East Sea where the varietygrows. Snow load also acted as an important driving force for structural modifications of A. japonica var. borealis fromcellular level in leaves to the organization of branch extension growth. Global warming by changing snowfall patterns inJapan may lead to range shifts in the two varieties of A. japonica

Distribution of Aucuba japonica in two contrasting geobotanical regions of Japan: An analysis of adaptation mode