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Cave-obligate species richness is related with habitat availability, namely the number of caves (Christman & Culver, 2001; Culver et al., 2003, 2004), but it is usually low compared with the richness of surface habitats for most taxa. Caves are usually restricted to certain geological areas, such as karstic regions and volcanic areas and these can be seen as islands surrounded by inhospitable habitat. Disjunction of cave systems and consequent reduced dispersal of organisms are important in determining that cave-obligate species usually exhibit narrower ranges and higher endemics proportions than their epigean counterparts (Barr & Holsinger, 1985; Culver et al., 2000; Sharratt et al., 2000; Gibert &

Diversity and community assembly patterns of epigean vs. troglobiont spiders in the Iberian Peninsula