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College of Pharmacy, Yeungnam University, Gyeongsan 712-749, Korea College of Pharmacy, Catholic University of Daegu, Gyeongsan 712-702, Korea. E-mail: woomh@cu.ac.kr College of Natural Science, Yeungnam University, Gyeongsan 712-749, Korea College of Pharmacy, Chungbuk National University, Cheongju 361-763, Korea College of Life Science, Catholic University of Daegu, Gyeongsan 712-702, Korea Received April 8, 2006

DNA Topoisomerases I and II Inhibition and Cytotoxicity of Constituents from the Roots of Rubia cordifolia