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The tommotiid Camenella reticulosa is redescribed based on new collections of well preserved sclerites from the Arrowie Basin (Flinders Ranges), South Australia, revealing new information concerning morphology and micro- structure. The acutely pyramidal mitral sclerite is described for the first time and the sellate sclerite is shown to be coiled through up to 1.5 whorls. Based on Camenella, a model is proposed by which tommotiid sclerites are composed of alternating dense phosphatic, and presumably originally organic-rich, laminae. Camenella is morphologically most similar to Lapworthella, Kennardia, and Dailyatia, and these taxa are interpreted to represent a monophyletic clade, here termed the camenellans, within the Tommotiida. Potential reconstructions of the scleritome of Camenella are discussed and although a tubular scleritome construction was recently demonstrated for the tommotiids Eccentrotheca and Paterimitra, a bilaterally symmetrical scleritome model with the sclerites arranged symmetrically on the dorsal surface of a vagrant animal can not be ruled out.16 page(s

The TommotiidCamenella reticulosafrom the Early Cambrian of South Australia: Morphology, Scleritome Reconstruction, and Phylogeny