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The family Equidae includes a diversity of extant species in the single genus Equus, notably horses (E. caballus and E. przewalskii) and a group called the non-caballines (which includes both Asian and African asses, as well as African zebras). The fossil record, however, shows even more remarkable diversity : within the past 5 million years at least 12 equid genera have been documented, including Hipparion, which is known from Plio-Pleistocene deposits in the Cradle of Humankind World Heritage Site in South Africa. A linear evolutionary sequence from a hyracotherian ancestor was proposed by Othaniel Marsh and championed by Thomas Huxley, but the evolution of equids within the past 55 million years is now recognized as being much more complex. Palaeontologists such as Basil Cooke, Rufus Churcher, Vera Eisenmann and Alan Gentry have generally relied on morphology and measurements for purposes of addressing the taxonomy of southern African Equidae, as others have done for equids from other areas. More recently the potential of genetic data has been demonstrated, most remarkably in a study published in December by Orlando et al.

Ancient DNA from fossil equids: A milestone in palaeogenetics