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Early late Pliocene rodent tooth remains, have been found in situ within Quercy paleokarstic fillings. They provide evidence of a sedimentary episode related to a high Pliocene marine level ca 3.5 Ma ago, that illustrates the situation predating the regional erosion phase and setting up of the hydrographic system.

Early Late Pliocene Paleokarstic Fillings Predating the Major Plio-Pleistocene Erosion of the Quercy Table, SW-France