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This paper is the first comprehensive work on the pupae of   Staphylinidae  . It is the first-ever attempt to employ the morphological characters of these pupae in phylogenetic analysis. The study shows that the external structures of   Staphylinini   pupae could be a useful, alternative source for assessing the relationships of certain taxa within a tribe. The paper also includes an illustrated key to the identification of pupae at the subtribe and generic levels (   Abemus   ,    Acylophorus   ,    Astrapaeus   ,    Atanygnathus   ,    Bisnius   ,    Creophilus   ,    Emus   ,    Erichsonius   ,    Gabrius   ,    Hesperus   ,    Heterothops   ,    Neobisnius   ,    Ocypus   ,    Ontholestes   ,    Philonthus   ,    Quedius   ,    Quedionuchus   ,    Rabigus   ,    Staphylinus   , and    Tasgius   ) of the tribe   Staphylinini  , found in Europe. Based on current knowledge of the morphology of pupal stages of   Staphylinini   species, eight morphological pupal types are presented:   Acylophorus  ,   Astrapaeus  ,   Atanygnathus  ,   Erichsonius  ,   Heterothops  ,   Philonthus  ,   Quedius   and   Staphylinus  . The paper also comments on pupal habitat, phenology and morphology in the context of antipredator and environmental adaptations.

Pupae of the mega-diverse rove beetle tribe Staphylinini ( Coleoptera , Staphylinidae ): their traits and systematic significance