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The Tailed Punch,    Dodona   eugenes   , is widely distributed in East Asia with seven subspecies currently recognized. However, two of them, namely ssp.formosana   and ssp.esakii   found in Taiwan, are hard to distinguish from each other due to ambiguous diagnostic characters. In this study, their taxonomic status is clarified by comparing genitalia characters and phylogenetic relationships based on mitochondrial sequences,  COI  and  COII  (total 2211 bps). Our results show that there is no reliable feature to separate these two subspecies. Surprisingly we found that    Dodona    in Taiwan is more closely related to the Orange Punch,    D.   egeon   , than to other subspecies of    D.   eugenes   . Therefore, the following nomenclatural changes are proposed:    Dodona   eugenes   formosana    is revised to specific status as    Dodona   formosana    Matsumura, 1919,  stat. rev , and ssp.esakii   is sunk to a junior synonym of    Dodona   formosana    syn. n.

zookeys

A distinct species,    Dodona   formosana   , detected in the    Dodona   eugenes    species complex: clarification of the taxonomic status of the Punch butterfly in Taiwan