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The    Thereus   oppia    species group includes species with and without a scent pad, which is a histologically and morphologically characterized male secondary sexual structure on the dorsal surface of the forewing. To assess the hypothesis that these structures are lost evolutionarily, but not regained (Dollo’s Law), the taxonomy of this species group is revised.    Thereus   lomalarga    sp. n. , and    Thereus   brocki    sp. n. , are described. Diagnostic traits, especially male secondary structures, within the    T.   oppia    species group are illustrated. Distributional and biological information is summarized for each species. Three species have been reared, and the caterpillars eat   Loranthaceae  . An inferred phylogeny is consistent with the hypothesis that scent pads in the    T.   oppia    species group have been lost evolutionarily twice (in allopatry), and not re-gained.

Male secondary sexual structures and the systematics of the Thereus oppia species group ( Lepidoptera , Lycaenidae , Eumaeini )