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Water beetles of the families   Gyrinidae  ,   Haliplidae  ,   Noteridae  , and   Dytiscidae   (aquatic   Adephaga  ) of the Makay in central-western Madagascar were surveyed in three campaigns during the years 2016–2018. A total of 74 species was collected from 62 sampling sites, all except one being newly recorded from the Makay.    Copelatus   malavergnorum    sp. nov.  (irinus    group) and    C.   zanabato    sp. nov.  (erichsonii    group) (  Dytiscidae  ,   Copelatinae  ) are described and their habitus and male genitalia are illustrated. A systematic account is given, including description of habitat preferences for each species. Analyses of species composition and dominance, species diversity and endemism highlighted the strong singularity of the aquatic   Adephaga   fauna inhabiting the sandstone massif of inner Makay (notably with several local endemic dytiscids) with respect to its peripheral lowlands. These comparisons were also performed between groups of sites categorised according to vegetation context (forested, semi-forested, non-forested). Rather unexpectedly, inner Makay although well-preserved and little deforested has relatively low endemism level and low species diversity (H 1  Hill number twice lower than in the geographically close and geologically similar massif of Isalo). Species diversity was higher in the deforested and man-impacted peripheral sites, which yielded a rich contingent of western Madagascar lowland species including a few undescribed or rarely observed dytiscids.

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﻿The aquatic   Adephaga   of the Makay, central-western Madagascar, with description of two new diving beetle species (  Coleoptera  ,   Gyrinidae  ,   Haliplidae  ,   Noteridae  ,   Dytiscidae  )

﻿The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species ( Coleoptera , Gyrinidae , Haliplidae , Noteridae , Dytiscidae )

The aquatic Adephaga of the Makay, central-western Madagascar, with description of two new diving beetle species ( Coleoptera , Gyrinidae , Haliplidae , Noteridae , Dytiscidae )