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Five Hawaiian species of    Paratachys    Casey are revised, including four newly described:    Paratachys   terryli    from Kauai;    P.   perkinsi    from Moloka‘i;    P.   haleakalae    from Maui; and    P.   aaa    from Hawai‘i Island. A lectotype is designated for the fifth Hawaiian species currently combined with    Paratachys   ,    Tachys   arcanicola    Blackburn, 1878 of Oahu. Hawaiian    Paratachys    spp. known from more than one specimen exhibit some degree of ocular polymorphism, that variation being extreme in    P.   terryli    where individuals range in ocular development from macrophthalmic with broadly convex eyes to microphthalmic with small, flat eyes. All Hawaiian    Paratachys    species comprise individuals with vestigial wings, with the exception of    P.   terryli   , where a single macropterous, macrophthalmic female complements the other 18 brachypterous specimens. Based on a transformation series of characters from the male aedeagus, the biogeographic history of Hawaiian    Paratachys    is consistent with progressive colonization of the Hawaiian Island chain. Three of the species do not appear to represent species of conservation concern, with    P.   terryli    and    P.   haleakalae    known from terrestrial deep soil, litter, and streamside microhabitats in montane wet rain forest, and the troglobitic    P.   aaa    occupying the dark zone of numerous, recently developed lava tube caves within the Mauna Loa and Kilauea volcanic massifs. The conservation status of the other two species is much more dire, with    P.   arcanicola    of O‘ahu not seen in nature since the early 20 th  Century, and    P.   perkinsi    known only from a single specimen fortuitously collected in 1894 near sea level on Moloka‘i.

Hawaiian Paratachys Casey ( Coleoptera , Carabidae ): small beetles of sodden summits, stony streams, and stygian voids