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.We sequenced multiple genes from the enigmatic genera    Bhagadatta    Moore, 1898,    Kumothales    Overlaet, 1940 and    Harmilla    Aurivillius, 1892 (  Nymphalidae  ,   Limenitidinae  ) and analysed them together with a large published dataset. We find that    Bhagadatta    is sister to the genera    Cymothoe    Hübner, 1819+   Harma    Doubleday, 1848, and that    Kumothales    is sister to these three.    Harmilla    is nested within the genus    Euriphene    Boisduval, 1847. We thus transfer    Kumothales    and    Bhagadatta    to the tribe    Cymothoini   , and we synonymise    Harmilla    syn. nov.  with    Euriphene   .

nota lepidopterologica

The phylogenetic positions of    Bhagadatta    Moore, 1898,    Kumothales    Overlaet, 1940 and    Harmilla    Aurivillius, 1892 (  Lepidoptera  ,   Nymphalidae  ,   Limenitidinae  ) based on molecular data