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DNA sequences from the nuclear LSU and ITS regions were used for phylogenetic analyses of   Thelephorales   with a focus on the stipitate hydnoid genera    Hydnellum    and    Sarcodon   . Analyses showed that    Hydnellum    and    Sarcodon    are distinct genera but that the current division, based on basidioma texture, makes    Sarcodon    paraphyletic with respect to    Hydnellum   . In order to make genera monophyletic several species are moved from    Sarcodon    to    Hydnellum    and the following new combinations are made:    Hydnellum   amygdaliolens   ,    H.   fennicum   ,    H.   fuligineoviolaceum   ,    H.   fuscoindicum   ,    H.   glaucopus   ,    H.   joeides   ,    H.   lepidum   ,    H.   lundellii   ,    H.   martioflavum   ,    H.   scabrosum   ,    H.   underwoodii   , and    H.   versipelle   . Basidiospore size seems to separate the genera in most cases.    Hydnellum    species have basidiospore lengths in the range 4.45−6.95 µm while the corresponding range for    Sarcodon    is 7.4−9 µm.    S.   quercinofibulatus    deviates from this pattern with an average spore length around 6 µm. Neotropical    Sarcodon    species represent a separate evolutionary lineage.

Reassessment of the generic limits for Hydnellum and Sarcodon ( Thelephorales , Basidiomycota )