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The most prolific source of exquisitely preserved bryophyte fossils is amber, which often contains inclusions in a three-dimensional and life-like state of preservation. In recent years, many fossil species have been described from 16 worldwide deposits ranging in age from the Miocene to the Cretaceous. One of the oldest is mid-Cretaceous Kachin amber from Myanmar. It includes the moss genera    Vetiplanaxis    and    Calymperites    as well as the leafy liverwort genera    Frullania   ,    Gackstroemia   ,    Protofrullania   , and    Radula   . All liverwort fossils belong to the mainly epiphytic   Porellales  , a group which was probably strongly influenced by the rise of angiosperms and underwent significant lineage turnover in the Cretaceous. Hence, Kachin amber provides important information on the evolution of leafy liverworts during a period characterized by fundamental changes in terrestrial ecosystems. Here, we describe two new species of the mainly epiphytic leafy liverwort genus    Radula    (  Radulaceae  ),    R.   patrickmuelleri    sp. nov.  and    R.   tanaiensis    sp. nov. , and emend the description of    Frullania   kachinensis    (  Frullaniaceae  ).    Radula    is now represented by four species and one fossil only described to genus level, which indicates a high diversity already in the mid-Cretaceous. Furthermore, we describe additional fossil specimens of   Frullaniaceae  , including the third fertile specimen of    Frullania   baerlocheri   , one sterile fossil of    F.   cretacea   , and 12 sterile fossils of    Protofrullania   cornigera   . The new fossil evidence necessitates an emendation of a recently published determination key for Cretaceous   Jungermanniidae  .

fossil record

﻿Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of    Radula   patrickmuelleri   , sp. nov. and    R.   tanaiensis   , sp. nov. (  Jungermanniopsida  ,   Porellales  ,   Radulaceae  )

﻿Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri , sp. nov. and R. tanaiensis , sp. nov. ( Jungermanniopsida , Porellales , Radulaceae )

Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri , sp. nov. and R. tanaiensis , sp. nov. ( Jungermanniopsida , Porellales , Radulaceae )