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Beetles previously considered to be    Ocys   harpaloides    (Audinet-Serville) from northern Africa, Spain, France, the United Kingdom, France, and Belgium belong to two species. These species can be distinguished using DNA sequences of  28S  rDNA, 18S rDNA,  COI ,  CAD , and topoisomerase I. A key, diagnoses, and images are provided to enable identification of specimens based upon characteristics of male and female genitalia, as well as microsculpture and other external structures. Through examination of the holotype of   Bembidium   harpaloides   v.   tachysoides   Antoine, as well as designation of lectotypes of    Bembidion   harpaloides    Audinet-Serville and    Ocys   melanocephalus    Stephens, and designation of a neotype for    Tachis   rufescens    Guérin-Ménéville, the valid names of the two species were determined to be    Ocys   harpaloides    and    Ocys   tachysoides    (Antoine).

deutsche entomologische zeitschrift

Hidden species within the genus    Ocys    Stephens: the widespread species    O.   harpaloides    (Audinet-Serville) and    O.   tachysoides    (Antoine) (  Coleoptera  ,   Carabidae  ,   Bembidiini  )