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The species of the    Melitaea   ala    Staudinger, 1881 complex are distributed in Central Asia. Here we show that this complex is a monophyletic group including the species,    M.   ala   ,    M.   kotshubeji    Sheljuzhko, 1929 and    M.   enarea    Fruhstorfer, 1917. The haploid chromosome number n=29 is found in    M.   ala    and    M.   kotshubeji    and is, most likely, a symplesiomorphy of the    M.   ala    complex. We show that    M.   ala    consists of four subspecies:    M.   ala   zaisana    Lukhtanov, 1999 (=   M.   ala   irtyshica    Lukhtanov, 1999,  syn. nov. ) (South Altai, Zaisan Lake valley),    M.   ala   ala    (Dzhungarian Alatau),    M.   ala   bicolor    Seitz, 1908 (North, East, Central and West Tian-Shan) and    M.   ala   determinata    Bryk, 1940 (described from “Fu-Shu-Shi”, China). We demonstrate that    M.   kotshubeji   kotshubeji    (Peter the Great Mts in Tajikistan) and    M.   kotshubeji   bundeli    Kolesnichenko, 1999 (Alai Mts in Tajikistan and Kyrgyzstan) are distinct taxa despite their geographic proximity in East Tajikistan.    Melitaea   enarea    is widely distributed in the southern part of Central Asia and is sympatric with    M.   kotshubeji   .

Chromosomal and DNA barcode analysis of the Melitaea ala Staudinger, 1881 species complex ( Lepidoptera , Nymphalidae )