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Coenosiini   (  Diptera  :   Muscidae  ) is a large cosmopolitan tribe of muscids, within which    Coenosia    Meigen and    Neodexiopsis    Malloch are the genera with the largest number of species. In this work, we revised for the first time, all the species placed by  Malloch (1934)  under    Austrocoenosia   , an endemic genus from the Andean Patagonian Forests, whose species are now placed in    Coenosia    and    Neodexiopsis   . We provide detail redescriptions for eight species placed by  Malloch (1934)  under    Austrocoenosia   , and decribed two new species (   Coenosia   delneneo    sp. nov.  and    Coenosia   patagonica    sp. nov. ), with high quality photographs detailing new structures of the male and female terminalia. To establish the position of the species of    Austrocoenosia    with respect to    Coenosia    and    Neodexiopsis   , we made a phylogenetic analysis using implied weighting for 36 taxa and 132 morphological characters of adults, including male and female genitalia. We recovered all species of    Austrocoenosia    as    Coenosia    species. We propose the following nomenclature actions:    Austrocoenosia    as a junior synonymy of    Coenosia    ( syn. rest. );    Coenosia   brevicornis    (Malloch) ( comb. nov. ),    Coenosia   dubia    (Bigot) ( comb. rest. );    Coenosia   hucketti   , Pont ( nom. nov. ) and    Coenosia   nigerrima    (Malloch) ( comb. rest. ). We also propose    Coenosia   spumicola    Pont as an unplaced species of    Coenosia    sensu lato. Finally, we updated the geographic distributions for all species and observed several new male and female terminalia structures, which enriched the discussion of the genera and the tribe.

arthropod systematics andamp; phylogeny

Phylogeny of the old and fragmented genus    Austrocoenosia    Malloch reveals new evidences on the morphology and evolution of the genera    Coenosia    Meigen and    Neodexiopsis    Malloch (  Diptera  :   Muscidae  )