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Recluse or violin spiders in the genus    Loxosceles    (  Scytodoidea  :   Sicariidae  ) are a diverse group (~140 extant species) including medically important species and distributed mainly in the Americas, Africa, and the Mediterranean region. In addition, this genus includes three fossil species from Miocene Dominican amber. Here we revise the taxonomy of these fossil species by examining, imaging and re-describing their type specimens. We find that    L.   defecta    Wunderlich, 1988 and    L.   deformis    Wunderlich, 1988 are  bona fide  members of the genus and report additional characters overlooked in their original descriptions. We further study the holotype of    L.   aculicaput    Wunderlich, 2004 using synchrotron radiation micro-computed tomography to reveal previously unknown morphological details hidden by fissures in the amber. We found several characters inconsistent with    Loxosceles    but consistent with    Drymusa    (false violin spiders;   Scytodoidea  :   Drymusidae  ), such as three claws, well-developed podotarsite, and a broad colulus. This suggests the species is misplaced in    Loxosceles   . To test this hypothesis, we estimated a total-evidence phylogeny of the superfamily   Scytodoidea   including extant and fossil taxa, morphological data, traditional molecular markers, and sequences of ultra-conserved elements. The results show unambiguously that    L.   aculicaput    belongs to    Drymusa    and is a close relative of extant species of the genus inhabiting the Greater Antilles. Therefore, we here transfer this species to    Drymusa   , establishing a new combination and new family assignment.    Drymusa   aculicaput    comb. nov.  represents the first known fossil   Drymusidae   and shows that crown members of this genus already existed in the Miocene.

arthropod systematics andamp; phylogeny

Revision of recluse spiders (  Araneae  :   Sicariidae  :    Loxosceles   ) preserved in Dominican amber and a total-evidence phylogeny of   Scytodoidea   reveal the first fossil   Drymusidae