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The weevil genus    Typoderus    is for the first time reported west of the Congo basin. Analysis of 2,136 aligned positions from one mitochondrial and two nuclear fragments revealed a moderately supported clade of five new Cameroonian species:    T.   amphion    sp. nov.  (Mt. Oku),    T.   canthus    sp. nov . (Mt. Oku),    T.   clytius    sp. nov.  (Mt. Cameroon),    T.   iphitus    sp. nov.  (Mt. Kupe) and    T.   telamon    sp. nov.  (Mt. Kupe). Molecular clock analysis of 20 DNA barcode fragments using a fixed substitution rate estimated divergences within this clade to be during the Middle to Late Miocene (10.5–5.4 million years ago,  MYA ), which pre-dates the onset of the Pliocene-Pleistocene global climatic fluctuations and corresponding cycles of African forest size fluctuation. Such relatively old dates are unexpected and might reflect four unavoidable shortcomings of the temporal analysis: 1. undersampled ingroup, 2. scarcity of comparative temporal data for other animal clades from the Cameroon Volcanic Line, 3. oversimplification of a fixed-rate molecular clock approach using a single maternally-inherited protein-coding marker and 4. possible overestimation of comparatively old ages when using largely saturated mitochondrial sequences. Two obscure weevil species from the Republic of the Congo are hypothesized to belong to the genus    Typoderus   :    T.   distinctus    (Hoffmann, 1968)  comb. nov.  (from   Anchonidium   subgenus   Neoanchonidium  ) and    T.   baloghi    (Hoffmann, 1968)  comb. nov.  (from   Anchonidium   subgenus   Subanchonidium  ). Three genus-group names are newly synonymized under    Typoderus   :Entypoderus    Voss, 1965  syn. nov.  (the only non-nominative subgenus of    Typoderus   ),Neoanchonidium    Hoffmann, 1968  syn. nov.  (subgenus of    Anchonidium   ) and    Subanchonidium    Hoffmann, 1968  syn. nov.  (subgenus of    Anchonidium   ). Habitus images and other supplementary information of all sequenced specimens are available online at  dx.doi.org/10.5883/DS-VGDS005  and  dx.doi.org/10.5883/DS-VGDS006 .

arthropod systematics andamp; phylogeny

Sky islands of the Cameroon Volcanic Line support the westernmost clade of five new    Typoderus    weevils (  Coleoptera  :   Curculionidae  :   Molytinae  )