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The family   Polymitarcyidae  , with a worldwide distribution, includes mayflies with large burrowing nymphs. South America harbors the highest diversity of this family, including the subfamilies   Asthenopodinae   and   Campsurinae  . In this work, three new species belonging to the genera    Tortopsis    and    Tortopus    (  Campsurinae  ) are described based on adults and nymphs from Colombia:    Tortopsis   toro    sp. nov. ,    Tortopsis   andaki    sp. nov.  and    Tortopus   coreguaje    sp. nov.  Additionally,    Tortopsis   limoncocha    is firstly recorded from Colombia. A cladistic analysis of all the species in these genera is presented, using external morphological characters of adults and eggs. Keys to male and female adults of all the species of both genera are presented.

arthropod systematics andamp; phylogeny

New insights into the phylogeny of    Tortopus    Needham and Murphy and    Tortopsis    Molineri (  Ephemeroptera  ,   Polymitarcyidae  ) with description of three new species