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Late Cretaceous shallow marine mollusc assemblages show a distinct vertical differentiation into those of the inner and the outer shelf. The realms Theia (pro Tethys) and Kalais (pro Boreal, Temperate) can only be distinguished by means of inner shelf assemblages. Outer shelf assemblages are uniform across realm boundaries. The boundary between the inner and the outer shelf is biologically significant. It is interpreted to reflect the lower boundary of the tropical/subtropical surface mixed layer and extended as far north as southern Sweden in the Late Cretaceous.

Marine Palaeobiogeography of the Central European Late Cretaceous