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We synthesise the generic properties of ecologically realistic multi-trophic level models and define criteria for ecological realism. We define three simple properties that all sensible ecosystem models should have: 1. Independence of scale: the functions that describe the change in the size of the population ( x i ) are independent of the scale at which we consider the population, that is: 1 xi dx i dt = f i x 1 , x 2 ,, x n ; N () for i = 1,2,,n . (1)

Consistency, competitive exclusion and coexistence in complex plankton ecosystem models