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Simple Summary  The mid-Cretaceous amber of Myanmar (also known as Burmese amber) is almost 100 MYA old and represents an invaluable source of information about the evolution of life in the late Mesozoic. This is particularly true for the early flies (lower Diptera) which underwent rapid radiation during the Cretaceous period. Here we describe a remarkable nematocerous fossil fly which shows a character typical of the flies of the suborder Brachycera—with strongly modified antenna—representing new evidence that such a Brachycera-like modification of the antennae has evolved several times during the evolutionary history of Diptera.    Abstract  A new fossil genus of Bibionidae (Diptera: Bibionomorpha),  Burmahesperinus  gen. nov., from the mid-Cretaceous Burmese amber, is described and illustrated (type species  Burmahesperinus antennatus  sp. nov., the other two species included are  B. conicus  sp. nov. and  B. pedicellatus  sp. nov.). The new genus is tentatively placed in a new subfamily, Burmahesperininae subfam. nov. of the family Bibionidae. Its possible phylogenetic position is briefly discussed. The new genus, as well as the subfamily, possesses the wing venation similar to the recent genus  Hesperinus  Walker, 1848, in combination with Brachycera-like modification of both the male and female antenna and the overall habitus typical of fungus gnats (Sciaroidea).

A Peculiar New Genus of Bibionomorpha (Diptera) with Brachycera-Like Modification of Antennae from Mid-Cretaceous Amber of Myanmar