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During the last decade, intracellular actin waves have attracted muchattention due to their essential role in various cellular functions, rangingfrom motility to cytokinesis. Experimental methods have advanced significantlyand can capture the dynamics of actin waves over a large range ofspatio-temporal scales. However, the corresponding coarse-grained theory mostlyavoids the full complexity of this multi-scale phenomenon. In this perspective,we focus on a minimal continuum model of activator-inhibitor type and highlightthe qualitative role of mass-conservation, which is typically overlooked.Specifically, our interest is to connect between the mathematical mechanisms ofpattern formation in the presence of a large-scale mode, due tomass-conservation, and distinct behaviors of actin waves.

Why A Large Scale Mode Can Be Essential For Understanding Intracellular Actin Waves