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A fundamental goal of systems neuroscience is to probe the dynamics of neuralactivity that drive behavior. Here we present an instrument to simultaneouslymanipulate neural activity via Channelrhodopsin, monitor neural response viaGCaMP3, and observe behavior in freely moving C. elegans. We use the instrumentto directly observe the relation between sensory stimuli, interneuron activityand locomotion in the mechanosensory circuit.

Simultaneous optogenetic manipulation and calcium imaging in freely moving C. elegans